In a standard installation you can customize this by creating a community pr. folder (i.e. at the toplevel). Then create a section pr. community (folder) , assign these sections to the different folders/communities. In this way, you can also customize the design pr. community-template (using the section_id).
Regarding access, you can assign a role with rights pr. section, and create a usergroup connected to the role where the users are given access to the specific community they are attached to.
In the user registerprocess, you have to make sure that the users are registered in the correct usergroup, then you're almost done ;)
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